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Vol. 36(2), 2001

could be considered as a subfamily of Potamididae.

In comparison with the data on Cerithidea califomica given by Bright (1958; 1960), C. costata differs in having esophageal crop absent, duet to digestive gland inserting in stornach (and not in distal extremity of esophagus), opened pallia) oviduct and bursa copulatrix present. Driscoll (1972) presents a detailed study on digestive system of C. californica, in which the stornach of C. costata, in particular, is very similar, differs in lacking the ridges near intestine origin.

Family Campanilidac Genus Campanile Fischer, 1884 (iype species: Cerithium giganteum Lamarck,

1804 [Eocene])

Campanile symbolicum lredale, 1917 (Figs 42, 81, 111,406-422)

Synonymy see Houbrick (198la: 282). Complement:

Campanile symbolicum; Abbott & Dance, 1983: 68 (fig.).

Description.

Shell (fig. 42). Relatively large (morę than 240 mm), tumform, heavy, thick walled. Whorls with almost straight outer surface, suture almost invisible, Periostracum white, calcified. Other details in Houbrick (198 la: 266-268, figs. 1-2).

Head-foot (figs 406, 408, 411). Homogeneous beige in color. Head protruding. Snout relatively smali, cylindrical, with a deep anterior, ventral, longitudinal, furrow. Tentacles somewhat long, broad, dorso-ventrally flattened. Eyes dark, on smali ommatophores situated approximately in middle-outer region of tentacles. Foot laige (about 1/3 whorl); fuirow of pedał glands deep, with thick borders in anterior margin, gradually becomes narrow in lateral region, surrounds en tire foot sole. Columellar muscle long, about 2.5 whorls, flattened and thick. Other details in Houbrick (1981a: 268-271, figs. 3a, b, 4a). Dorsal wali of haemocoel thick muscular in anterior half; posterior half thinner; separating both regions a diaphragm-like septum surrounding esophagus (fig. 411: o 1).

Operculum. Sub-circular, comeus, sub-cen-tral nucleus, palcispiral; occupies paitially aperture (fig.408) (see also Houbrick, 198la: 268, figs le-f).

Mantle organs (figs 407, 410, 412, 422). Mantle border entirely with smali papillae, somewhat uniform in size; in region of siphonal canal a deep concavity. Mantle cavity very deep, morę than 2 whorls. Osphradium long, bipectinate, somewhat elliptical outline; length little morę than 1/3 of gili length; filaments very thin and close with cach other, right filaments larger and discretely morę numerous than left filaments; right filaments attached to ctenidial vein and probably irrigating to it; left margin free; dorsal surface of osphradium altogether attached to mantle (fig. 422). Gili long, little morę than half of cavity, narrow, filaments triangular tali, tip pointed, almost central; gili begins relatively posterior fforn mantle border. Ctenidial vein broad sińce anterior gili margin, about 1/3 of its length (posterior) ffee from gili, running at left in fioor of pallial cavity, close kidney. Between gili and rectum a relatively broad space; in posterior 2/3 of this space several transverse vessels, sometimes dichotomic, inserted in an inconspicuous ad-rectal sinus. Hypobranchial glands a lamina covering vessels between gili and rectum; some specimens bearing a tali fold lying anterior half of rectum. Rectum described below. Anus siphoned, very posterior from mantle border. Other details in Houbrick (1981 a: 271-274, figs 3c, 4a, b).

Circulatory and excretory systems (figs 407, 409, 410). Heart similar to those of preceding species, proportionally smali, anterior and posterior aortas not attached directly in stornach, but in a lobe of digestive gland. Kidney very long, morę than half whorl, somewhat flattened, brown in color. Anterior-pallial half of kidney almost solid, bearing several glandular septa radially disposed; nephrostome a slit near posterior limit of pallial cavity. Posterior-visceral half of kidney with two regions, that at left lying pericardium, almost solid, bearing several transversal glandular septa; that at right replete of several vessels close with each other, connected in a main vessel which runs in median region (fig. 409).

Digestive system (figs 407, 411-421). Mouth deeply situated in median-ventral furTow of snout, bordered by several inner folds. Buccal mass proportionally very large, situated almost entire posterior to snout. About half of buccal mass constituting by greatly developed jaws and



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